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Some of them are mother-wasps and some are workers, as with the tamer kind; but it is by observation of the tame wasps that one may learn the varied characteristics of the mothers and the workers. For in the case of the tame wasps also there are two kinds; one consists of leaders, who are called mothers, and the other of workers. The leaders are far larger and milder-tempered than the others. The workers do not live over into a second year, but all die when winter comes on; and this can be proved, for at the commencement of winter the workers become drowsy, and about the time of the winter solstice they are never seen at all. The leaders, the so-called mothers, are seen all through the winter, and live in holes underground; for men when ploughing or digging in winter have often come upon mother-wasps, but never upon workers. The mode of reproduction of wasps is as follows. At the approach of summer, when the leaders have found a sheltered spot, they take to moulding their combs, and construct the so-called sphecons,-little nests containing four cells or thereabouts, and in these are produced working-wasps but not mothers. When these are grown up, then they construct other larger combs upon the first, and then again in like manner others; so that by the close of autumn there are numerous large combs in which the leader, the so-called mother, engenders no longer working-wasps but mothers. These develop high up in the nest as large grubs, in cells that occur in groups of four or rather more, pretty much in the same way as we have seen the grubs of the king-bees to be produced in their cells. After the birth of the working-grubs in the cells, the leaders do nothing and the workers have to supply them with nourishment; and this is inferred from the fact that the leaders (of the working-wasps) no longer fly out at this time, but rest quietly indoors. Whether the leaders of last year after engendering new leaders are killed by the new brood, and whether this occurs invariably or whether they can live for a longer time, has not been ascertained by actual observation; neither can we speak with certainty, as from observation, as to the age attained by the mother-wasp or by the wild wasps, or as to any other similar phenomenon. The mother-wasp is broad and heavy, fatter and larger than the ordinary wasp, and from its weight not very strong on the wing; these wasps cannot fly far, and for this reason they always rest inside the nest, building and managing its indoor arrangements. The so-called mother-wasps are found in most of the nests; it is a matter of doubt whether or no they are provided with stings; in all probability, like the king-bees, they have stings, but never protrude them for offence. Of the ordinary wasps some are destitute of stings, like the drone-bees, and some are provided with them. Those unprovided therewith are smaller and less spirited and never fight, while the others are big and courageous; and these latter, by some, are called males, and the stingless, females. At the approach of winter many of the wasps that have stings appear to lose them; but we have never met an eyewitness of this phenomenon.
Wasps are more abundant in times of drought and in wild localities.
They live underground; their combs they mould out of chips and earth, each comb from a single origin, like a kind of root. They feed on certain flowers and fruits, but for the most part on animal food. Some of the tame wasps have been observed when sexually united, but it was not determined whether both, or neither, had stings, or whether one had a sting and the other had not; wild wasps have been seen under similar circumstances, when one was seen to have a sting but the case of the other was left undetermined. The wasp-grub does not appear to come into existence by parturition, for at the outset the grub is too big to be the offspring of a wasp. If you take a wasp by the feet and let him buzz with the vibration of his wings, wasps that have no stings will fly toward it, and wasps that have stings will not; from which fact it is inferred by some that one set are males and the other females. In holes in the ground in winter-time wasps are found, some with stings, and some without.
Some build cells, small and few in number; others build many and large ones. The so-called mothers are caught at the change of season, mostly on elm-trees, while gathering a substance sticky and gumlike. Alarge number of mother-wasps are found when in the previous year wasps have been numerous and the weather rainy; they are captured in precipitous places, or in vertical clefts in the ground, and they all appear to be furnished with stings.
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So much for the habits of wasps.